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The mid-Holocene was an extremely dry period in the Lake Titicaca Basin of South America, when lake levels were at their lowest point in the Holocene. South of the lake, a lack of outflow and very low and irregular precipitation would have created desert-like conditions. This area's 'archaeological silence' seems to reflect an effective lack of population. This situation changed drastically as lake levels rose suddenly in the centuries following 3540 cal BP. As the desert disappeared, a flux of migrants filled the landscape, probably from the population concentration in the basin's western highlands. They imported and developed new technologies and economic practices and reorganized them into an agropastoral lifeway. The emergence of agropastoralism was both rapid and widespread, as people throughout the Lake Titicaca Basin adopted this practice. This major, regional shift can be productively framed as an adaptive cycle or Holling loop. This approach builds on the robust foundation of complexity theory, emphasizes the integrated nature of humans and their environment in a single system, highlights how systems fluctuate between slow and accelerated change, and is useful for developing hypotheses. Cascading feedback loops in climate, ecology, and cultural practices generated the emergence of agropastoralism. This resilient system is still in use today and is currently facing major climate changes, which makes understanding its origins especially relevant. (C) 2016 Elsevier Ltd and INQUA. All rights reserved.