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A new species narrowly endemic to the extremely arid western portion of the San Joaquin Desert, California, Navarretia panochensis, is here described. This species belongs to the N. pubescens group of Navarretia section Mitracarpium, sharing several features including the presence of three cotyledons. It is distinguished from these near relatives by having entire rather than forked cotyledons, generally lavender corolla lobes, and like some populations of N. pubescens, lacking a widened distal rachis of its upper leaves and bracts. A key distinguishing N. panochensis from other species of the N. pubescens group is provided.