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The analysis of organic-walled phytoplankton, pollen, and spores in sections of the Solenovian Horizon (second half of the lower Oligocene) in the Western Carpathians, Crimea-Caucasus region, and Scythian and Turan plates revealed that facies of a more desalinated basin (lower, Polbian Subhorizon) are characterized by a more impoverished assemblage of dinocysts, prasinophytes, and acritarchs numbering up to 20-30 taxa. Depending on facies properties, they are dominated by representatives of Batiacasphaera accompanied by thin-walled Hystrichokolpoma morphotypes or cysts of the Adnatosphaeridium-Glaphyrocysta Group, which allows facies to be defined as bed-ranked units within the single lower Oligocene Wetzeliella gochtii dinocyst zone. The organic-walled phytoplankton assemblage from the upper (Ikiburulian) subhorizon implies the frequently alternating salinity regime for this inner basin. The palynological assemblages of the Solenovian Horizon are characterized by a high share of Pinus pollen accompanied by pollen of Fagaceae (particularly in southern areas), Juglandaceae, and diverse thermophilic plants including evergreen taxa. The Solenovian climate was subtropical in western and southern areas and moderately warm seasonal with hot summers in others. The dominant arid climate was interrupted by brief humid episodes.