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During the last glacial period, a large part of the Aquitaine basin (southwest France) was a periglacial desert comprising coversands with low-relief dune fields surrounded by loess accumulations. OSL and radiocarbon dates show that the phase of maximum sand deposition coincides with Marine Isotope Stage (MIS) 2. Peats and gleyic palaeosoils intercalated within the sands at some sites indicate that vegetation cover was able to develop locally during short events, possibly D-O interstadials, due to raised groundwater levels in interdunal depressions. Few Palaeolithic sites have been discovered in the coversand area in contrast to the peripheral loess region. Systematic survey along a future highway corridor demonstrates that this paucity of sites is not the result of insufficient survey nor deep site burial, but rather reflects an archaeological reality. This strongly suggests that the sand area was not attractive for hunter-gatherer populations due to its reduced levels of water resources, and available vegetation and animal biomass. The distribution of cultural markers such as art items and projectile points also shows that the coversand area probably acted as a barrier separating two different cultural sub-areas, one in the Pyrenees and Cantabria, the other in the Perigord. As a consequence, the commonly accepted view that southwest France, as a whole, served as a refugia during the cold and arid phases of the Pleistocene should be replaced by a more complex one that reflects the fact that a large part of the territory was almost unoccupied and that human populations were concentrated along alluvial valleys. (C) 2013 Elsevier Ltd. All rights reserved.