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The Cal Basin formed in the late Miocene as an orogen-top rift hosting terrestrial sedimentation. The initial array of alluvial fans in a half-graben basin was replaced by an axial meandering-river system during the late Tortonian. Palaeomammal taxa indicate a mid-Turolian age of the deposits and a grass-dominated steppe ecosystem. Isotopic data from pedogenic carbonates indicate a warm, semiarid to arid climate. Subhumid to humid climatic conditions prevailed in the Pliocene, with a palustrine environment and savannah-type open ecosystem, recording a regional response to the marine flooding that terminated the Messinian ’salinity crisis’ in the Mediterranean. Pleistocene saw re-establishment of a fluvial system in the basin with the development of an open steppe ecosystem in warm, semiarid to arid climatic conditions. The sedimentary facies analysis of the basin-fill succession, combined with biostratigraphic data, render the basin a regional reference and help to refine the Neogene tectono-climatic history of SW Anatolia. (C) 2012 Academie des sciences. Published by Elsevier Masson SAS. All rights reserved.