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The global climate-vegetation model HadSM3_TRIFFID has been used to estimate the equilibrium states of climate and vegetation with pre-industrial and last glacial boundary conditions. The present study focuses oil the evaluation of the terrestrial biosphere component (TRIFFID) and its response to changes in climate and CO2 concentration. We also show how, by means of a diagnosis of the distribution of plant functional types according to climate parameters (soil temperature, winter temperature, growing-degree days, precipitation), it is possible to get better insights into the strengths and weaknesses of the biosphere model by reference to field knowledge of ecosystems.

The model exhibits profound changes between the vegetation distribution at the Last Glacial Maximum and today that are generally consistent with palaeoclimate data, such as the disappearance of the Siberian boreal forest (taiga), an increase in shrub cover in Europe and an increase of the Subtropical desert area. The effective equatorial and sub-tropical tree area is reduced by 18%. There is also an increase in cover of wooded species in North-Western Africa and in Mexico. The analysis of bioclimatic relationships turns out to be an efficient method to infer the contributions of different climatic factors to vegetation changes, both at high latitudes, where the position of the boreal treeline appears in this model to be more directly constrained by the water stress than by summer temperature, and in semi-humid areas where the contributions of temperature and precipitation changes may partly compensate each other. Our study also confirms the major contribution of the decrease in CO2 to environmental changes and carbon storage through its selective impact on gross primary productivity of C3 and C4 plants and a reduction by 25% of water-use efficiency. Specifically, the reduction in CO2 concentration increases the amount of precipitation necessary to sustain at least 20% of grass fraction by 50 mm/year; the corresponding threshold for trees is increased by about 150 mm/year. As a consequence, a reduction in CO2 concentration considerably widens the climatic range where grasses and shrubs dominate. Crown Copyright (c) 2004 Published by Elsevier B.V. All rights reserved.