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Barley (Hordeum vulgare) and wheat (Triticum aestivum) are similar crops that can be grown in the arid regions of southwestern Asia. Generally, barley is favored over wheat in drier areas. However, the quantitative basis for the relative advantage of each crop under differing growing conditions has not been resolved. This study was undertaken to compare the two crops using the same model structure. A wheat model developed for southwestern Asia was readily adapted to simulate barley. Simulations of barley crops proved to be robust over a range of environments. Comparison of simulated yields in a number of environments showed that the critical difference between species was the cumulative effect of plant development traits that allowed barley to mature much earlier than wheat. Consequently, early maturity of barley in contrast to wheat often allowed barley to complete its growing cycle before water-deficit conditions developed in the spring. There was, however, little or no yield advantage for barley in seasons of high rainfall. Also. rare seasons with significant rainfall after barley matured allowed wheat to actually produce the greater yield. These simulation results indicated that development traits are crucial in the interaction between crop yield and exposure to water deficits at the end of the growing season. Published by Elsevier B.V.