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The relationship between soil surface cryptogamic crusts and seed banks was investigated in the shrub-steppe in the Lower Columbia Basin. Seventy-four percent of the seeds in a disturbed bunchgrass community were found in crevices bordering cryptogamic crust polygons. In a sagebrush/bunchgrass community, 89% of the seeds were found in crevices. In a disturbed bunchgrass community, Bromus tectorum seeds were found in both the seed bank and aboveground vegetation communities. Bromus tectorum seeds were located in the seed bank of a sagebrush/bunchgrass community, although it bad a minor presence in the aboveground community. Seeds of Artemisia tridentata Nutt, were not found in either the bunchgrass or sagebrush/bunchgrass communities. The high number of seeds found in crevices bordering the cryptogamic crust suggests that crevices play a role in determining the small-scale distributional pattern of shrub-steppe plants at the Fitzner-Eberhardt Arid Lands Ecology Reserve.