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The biology, ecology, biogeography, phylogeny, and evolution of sand-dwelling scorpions are reviewed. We present general features of sand that form the ecological theatre on which psammophiles have evolved. Sand scorpions, compared to those that live off sand, are not very diverse in any one local habitat, although they are often relatively abundant. Worldwide, they are represented by 29 genera, 11 of which exist exclusively in sand. Endemic genera belong to non-related families, suggesting that psammophilic adaptations evolved in situ in different sand deserts of the world. (C) 1998 Academic Press.